Neuroendocrinology, neurobiology, morphological neuroscience of crustaceans and sea cucumbers
Aquaculture, reproductive biology, and endocrine physiology of crustaceans and sea cucumbers
Neurotransmitters, biogenic amines, and neuropeptides in regulation of reproduction and feeding of crustaceans and sea cucumbers
Brain morphology and neural circuits of decapod crustaceans (Macrobrachium rosenbergii and Litopenaeus vannamei), sea cucumbers (Holothuroidea) and other invertebrates